[1]      Yu S W, Zhu M Z, Li P, et al. Dissection of the spatial dynamics of biosynthesis, transport, and turnover of major amino acids in tea plants (Camellia sinensis) [J]. Horticulture Research, 2024, 11(5): 327-345.

[2]      Wei J C, Mu X Y, Wang S Y, et al. Integrated metabolome and transcriptome analysis provides insights into the mechanisms of terpenoid biosynthesis in tea plants (Camellia sinensis) [J]. Food Research International, 2024, 201(4): 115542-115563.

[3]      Yu J Y, Ho C T, Lin Z, et al. Sensomics-assisted characterization of key flowery aroma compounds in Lu'an Guapian green tea infusion (Camellia sinensis) [J]. Journal of Agricultural and Food Chemistry, 2023, 4(3): 486-501.

[4]      Qiao D H, Tang M S, Jin L, et al. A monoterpene synthase gene cluster of tea plant (Camellia sinensis) potentially involved in constitutive and herbivore-induced terpene formation [J]. Plant Physiology and Biochemistry, 2022, 184: 1-13. https://doi.org/10.1016/j.plaphy.2022.05.016.

[5]      Liu G H, Yang M, Fu J Y. Identification and characterization of two sesquiterpene synthase genes involved in volatile-mediated defense in tea plant (Camellia sinensis) [J]. Plant Physiology and Biochemistry, 2020, 155: 650-657. https://doi.org/10.1016/j.plaphy.2020.08.004.

[6]      Ida B N, Andari S, Satutya W, et al. Phytochemical screening, antioxidant activity, functional groups and chemical element characterization analysis of (-)-epigallocatechin-3-gallate (EGCG) in East Javanese green tea methanolic extract: an experimental in vitro study [J]. Systematic Reviews in Pharmacy, 2020, 11(5): 511-519.

[7]      Lin F M, Wu H N, Li Z L, et al. Effect of mechanical damage in green-making process on aroma of rougui tea [J]. Foods, 2024, 13(9): 1315. https://doi.org/10.3390/foods13091315.

[8]      Jing T T, Qian X N, Du W K, et al. Herbivore-induced volatiles influence moth preference by increasing the β-ocimene emission of neighbouring tea plants [J]. Plant, Cell & Environment, 2021, 44(11): 3667-3680.

[9]      Liu G H, Yang M, Yang X M, et al. Five TPSs are responsible for volatile terpenoid biosynthesis in Albizia julibrissin [J]. Journal of Plant Physiology, 2021, 259(258): 153358-153366.

[10]    Fauziah F, Permana A D, Faizal A. Characterization of volatile compounds from tea plants (Camellia sinensis (L.) Kuntze) and the effect of identified compounds on Empoasca flavescens behavior [J]. Horticulturae, 2022, 8(7): 623. https://doi.org/10.3390/horticulturae8070623.

[11]    Jiang D, Yang W H, Pi J H, et al. Genome-wide identification, characterization, and expression profiling of the glutaredoxin gene family in tea plant (Camellia sinensis) [J]. Forests, 2023, 14(8): 1647. https://doi.org/10.3390/f14081647.

[12]    Shen C, Li X. Genome-wide identification and expression pattern profiling of the ATP-binding cassette gene family in tea plant (Camellia sinensis) [J]. Plant Physiology and Biochemistry, 2023, 202(9): 107930-107942.

[13]    Chen F, Tholl D, D'Auria J C, et al. Biosynthesis and emission of terpenoid volatiles from Arabidopsis flowers [J]. The Plant Cell, 2003, 15(2): 481-494.

[14]    Cui J, Bahrami A K, Pringle E G, et al. Pseudomonas syringae manipulates systemic plant defenses against pathogens and herbivores [J]. PNAS, 2005, 102(5): 1791-1796.

[15]    Du B Y, Ma X L, Liu H, et al. Transcription factor MdLSD1 negatively regulates α-farnesene biosynthesis in apple-fruit skin tissue [J]. Plant Biology, 2022, 24(6): 1076-1083.

[16]    Wang X, Zeng L, Liao Y, et al. Formation of α-farnesene in tea (Camellia sinensis) leaves induced by herbivore-derived wounding and its effect on neighboring tea plants [J]. International Journal of Molecular Sciences, 2019, 20(17): 4151. https://doi.org/10.3390/ijms20174151.

[17]    Tian J H, Chen Z, Jiang C, et al. Chromosome-scale genome assembly of Docynia delavayi provides new insights into the α-farnesene biosynthesis [J]. International Journal of Biological Macromolecules, 2024, 278(3): 134820-134832.

[18]    Huang W J, Fang S M, Su Y L, et al. Insights into the mechanism of different withering methods on flavor formation of black tea based on target metabolomics and transcriptomics [J]. Food Science and Technology, 2023, 189(16): 115537-115549.

[19]    Zeng L, Jin S, Xu Y Q, et al. Exogenous stimulation-induced biosynthesis of volatile compounds: aroma formation of oolong tea at postharvest stage [J]. Critical Reviews in Food Science and Nutrition, 2022, 64(1): 76-86.

[20]    He C, Li Y, Zhou J, et al. Multi-omics and Co-expression network insights into aroma formation of suitable oolong tea cultivars during withering and shaking stage [J]. Food Research International, 2026, 233(1): 118971. https://doi.org/10.1016/j.foodres.2026.118971.

[21]    Zhang S, Xia H, Sun S, et al. Analysis of volatile metabolites and differences in aroma quality of green tea, black tea and oolong tea prepared from highly aromatic tea variety (Dancong) [J]. Food Research International, 2025, 221(3): 117402. https://doi.org/10.1016/j.foodres.2025.117402.

[22]    Wu S H, Yang Y H, Chen J M, et al. Histone deacetylase CsHDA6 mediates the regulated formation of the anti-insect metabolite α-farnesene in tea (Camellia sinensis) [J]. Plant Science, 2022, 326: 111501. https://doi.org/10.1016/j.plantsci.2022.111501.

[23]    Wei J C, Yang Y, Ye P, et al. Biosynthesis and the transcriptional regulation of terpenoids in tea plants (Camellia sinensis) [J]. International Journal of Molecular Sciences, 2023, 24(8): 6937-6948.

[24]    Zhao X M, Chen S, Wang S S, et al. Defensive responses of tea plants (Camellia sinensis) against tea green leafhopper attack: a multi-omics study [J]. Frontiers in Plant Science, 2020, 10(17): 1707-1721.

[25]    Mu D, Pan C, Qi Z, et al. Multivariate analysis of volatile profiles in tea plant infested by tea green leafhopper Empoasca onukii Matsuda [J]. Plant Growth Regulation, 2021, 95(1): 111-120.

[26]    Goyal L, Khanna S. Recent advances in microbial production of butanol as a biofuel [J]. International Journal of Applied Sciences and Biotechnology, 2019, 7(2): 130-152.

[27]    He J G, Feng Y X, Cheng Y D, et al. Changes in α-Farnesene and phenolic metabolism and the expression of associated genes during the development of superficial scald in two distinct pear cultivars [J]. International Journal of Molecular Sciences, 2022, 23(20): 12088-12101.

[28]    Zheng Y C, Hu Q C, Yang Y, et al. Architecture and dynamics of the wounding-induced gene regulatory network during the oolong tea manufacturing process (Camellia sinensis) [J]. Frontiers in Plant Science, 2022, 12(27): 788469-788483.

[29]    Ai Z Y, Hu S H, Ji L F, et al. The combination of shaking and yellow-light withering promote the volatile aroma components and the aroma quality of black tea [J]. Foods, 2025, 14(5): 758-772.

[30]    Pechous S W, Whitaker B D. Cloning and functional expression of an (E,E)-alpha-farnesene synthase cDNA from peel tissue of apple fruit [J]. Planta, 2004, 219(1): 84-94.

[31]    Wang Q, Liu H, Zhang M, et al. MdMYC2 and MdERF3 positively co-regulate α-farnesene biosynthesis in apple [J]. Frontiers in Plant Science, 2020, 11: 512844. https://doi.org/10.3389/fpls.2020.512844.

[32]    Gorina S S, Egorova A M, Lantsova N V, et al. Discovery of α-linolenic acid 16(S)-lipoxygenase: cucumber (Cucumis sativus L.) vegetative lipoxygenase 3 [J]. International Journal of Molecular Sciences, 2023, 24(16): 12977. https://doi.org/10.3390/ijms241612977.

[33]    Alicandri E, Paolacci A R, Osadolor S, et al. On the evolution and functional diversity of terpene synthases in the Pinus species: a review [J]. Journal of Molecular Evolution, 2020, 88(3): 253-283.

[34]    Tang X, Zhang X J, Pan J J, et al. Z/E configuration controlled by a Taxus sesquiterpene synthase facilitating the biosynthesis of (3Z,6E)-α-farnesene [J]. Phytochemistry, 2024, 229(3): 114304. https://doi.org/10.1016/j.phytochem.2024.114304.

[35]    Guo Z J, Zuo Y X, Wang S Y, et al. Early signaling enhance heat tolerance in Arabidopsis through modulating jasmonic acid synthesis mediated by HSFA2 [J]. International Journal of Biological Macromolecules, 2024, 267(Part 2): 131256. https://doi.org/10.1016/j.ijbiomac.2024.131256.

[36]    Schuman M C, Palmer-Young E C, Schmidt A, et al. Ectopic terpene synthase expression enhances sesquiterpene emission in Nicotiana attenuata without altering defense or development of transgenic plants or neighbors [J]. Plant Physiology, 2014, 166(2): 779-797.

[37]    Dao M, Zhang Z Y, Yin X, et al. CsAFS1 enhances the self-resistance of tea plants by promoting terpenoid accumulation [J]. Physiologia Plantarum, 2025, 177(4): e70430. https://doi.org/10.1111/ppl.70430.

[38]    Wang J Z, Dao M, Yang Z Y, et al. CsAFS2 gene from the tea plant intercropped with Chinese chestnut plays an important role in insect resistance and cold resistance [J]. Forests, 2024, 15(2): 380. https://doi.org/10.3390/f15020380.

[39]    Xu Q S, Zhuang Z N, Cheng L, et al. CsNAC29 transcription factor activates α-farnesene emission to resist gray blight disease in tea plant (Camellia sinensis) [J]. The Plant Journal, 2025, 123(3): e70421. https://doi.org/10.1111/tpj.70421.

[40]    Gu H L, Li J X, Qiao D H, et al. A defensive pathway from NAC and TCP transcription factors activates a BAHD acyltransferase for (Z)-3-hexenyl acetate biosynthesis to resist herbivore in tea plant (Camellia sinensis) [J]. New Phytologist, 2025, 245(3): 1232-1248.

[41]    Zeng L T, Liao Y Y, Li J L, et al. α-Farnesene and ocimene induce metabolite changes by volatile signaling in neighboring tea (Camellia sinensis) plants [J]. Plant Science, 2017, 264(3): 29-36.

[42]    Ban Y, Oyama-Okubo N, Honda C, et al. Emitted and endogenous volatiles in ‘Tsugaru’ apple: the mechanism of ester and (E,E)-α-farnesene accumulation [J]. Food Chemistry, 2010, 118(2): 272-277.

[43]    王茹茹, 邓帅, 丁瑞瑞, 等. 青香蕉苹果MdAFS基因序列与表达模式分析[J]. 生物技术通报, 2017, 33(7): 75-82.
Wang R R, Deng S, Ding R R, et al. The sequence and expression analysis of MdAFS gene in 'White Winter Pearmain' apple [J]. Biotechnology Bulletin, 2017, 33(7): 75-82.

[44]    苑克俊, 刘庆忠, 张力思, 等. 苹果α-法尼烯合酶基因的启动子序列及同源基因[J]. 基因组学与应用生物学, 2010, 29(5): 904-910.
Yuan K J, Liu Q Z, Zhang L S, et al. Promoter sequence and homologous genes of apple α-farnesene synthase gene [J]. Genomics and Applied Biology, 2010, 29(5): 904-910.

[45]    Portnoy V, Benyamini Y, Bar E, et al. The molecular and biochemical basis for varietal variation in sesquiterpene content in melon (Cucumis melo L.) rinds [J]. Plant Molecular Biology, 2008, 66(6): 647-661.

[46]    Varnika B, Jaya M, Ajay J, et al. Aphid-repellent pheromone E-β-farnesene is generated in transgenic Arabidopsis thaliana over-expressing farnesyl diphosphate synthase2 [J]. Annals of Botany, 2014, 115(4): 581-591.

[47]    Zhang X H, Chen X H, Jaime A, et al. Characterization of sandalwood (E,E)-α-farnesene synthase whose overexpression enhances cold tolerance through jasmonic acid biosynthesis and signaling in Arabidopsis [J]. Planta, 2023, 258(3): 54. https://doi.org/10.1007/s00425-023-04212-1.

[48]    Lin J Y, Wang D, Chen X L, et al. An (E,E)-α-farnesene synthase gene of soybean has a role in defence against nematodes and is involved in synthesizing insect-induced volatiles [J]. Plant Biotechnology Journal, 2016, 15(4): 510-519.

[49]    Song J, Lee G, Jung J, et al. Effect of soybean volatiles on the behavior of the bean bug, Riptortus pedestris [J]. Journal of Chemical Ecology, 2022, 48(2): 207-218.

[50]    Wang W, Wang M, Feng J, et al. Terpene synthase gene family in Chinese chestnut (Castanea mollissima BL.) harbors two sesquiterpene synthase genes implicated in defense against gall wasp Dryocosmus kuriphilus [J]. Journal of Agricultural and Food Chemistry, 2024, 72(3): 1571-1581.

[51]    Huang M S, Abel C, Sohrabi R, et al. Variation of herbivore-induced volatile terpenes among Arabidopsis ecotypes depends on allelic differences and subcellular targeting of two terpene synthases, TPS02 and TPS03 [J]. Plant Physiology, 2010;153(3):1293-1310.

[52]    Cao J J, Li D D, Wang X H, et al. UV-B and mechanical wounding synergistically induce α-farnesene biosynthesis via CsHY5-CsMYC2 cooperation during oolong tea processing [J]. Horticultural Plant Journal, 2026, 12(1): 161-171.

[53]    Gu D C, Wu S H, Yu Z M, et al. Involvement of histone deacetylase CsHDA2 in regulating (E)-nerolidol formation in tea (Camellia sinensis) exposed to tea green leafhopper infestation [J]. Horticulture Research, 2022, 9(28): uhac158. https://doi:10.1093/hr/uhac158.

[54]    Wang X Y, Ma H C, Jiang Q T, et al. The CsbHLH124-CsTPS5 regulatory module governs D-limonene biosynthesis and gray blight resistance in tea plant (Camellia sinensis) [J]. Plant, Cell & Environment, 2026, 49(2): 940-953.

[55]    Ye Z L, Zheng Z Q, Wang J S, et al. JA regulates caffeine biosynthesis in tea leaf for resistance against fungal infection and antagonises with GA to balance the defence-growth trade-off via the CsDELLA-JAZ-MYC2-MYB184-TCS1 module [J]. Plant Biotechnol Journal, 2025, 23(12): 5865-5890.

[56]    Li J X, Li M, Gu H L, et al. CsNAC2 and CsMYB306 orchestrate a jasmonate signaling cascade contributing to tea plant resistance to Ectropis obliqua [J]. Plant Physiology, 2025, 199(4): kiaf614. https://doi.org/10.1093/plphys/kiaf614.

[57]    Wang Z L, Yuan Y X, Dong R, et al. Salicylic acid regulates biosynthesis of floral fragrance (E)-β-farnesene via NPR3-WRKY1 module in chrysanthemum [J]. Molecular Horticulture, 2025, 5(1): 52. https://doi.org/10.1186/s43897-025-00174-y.

[58]    Huang J Y, Zhao X B, Bürger M, et al. The role of ethylene in plant temperature stress response [J]. Trends in Plant Science, 2023, 28(7): 808-824.

[59]    Huang C, Tang J W, Sun W Q, et al. Gibberellin modulates the post-transcriptional regulation of CsHDZ34 to positively regulate bud break in tea plants [J]. New Phytologist, 2026, 249(4): 1850-1870.

[60]    Gai Z S, Wang Y, Ding Y Q, et al. Exogenous abscisic acid induces the lipid and flavonoid metabolism of tea plants under drought stress [J]. Scientific Reports, 2020, 10(1): 12275. https://doi.org/10.1038/s41598-020-69080-1.

[61]    吴晴阳, 周子维, 武清扬, 等. 乌龙茶加工过程中α-法呢烯的形成关键调控基因的筛选与表达分析[J]. 食品工业科技, 2020, 41(15): 135-142.
Wu Q Y, Zhou Z Y, Wu Q Y, et al. Screening and expression analysis of key regulatory genes for α-farnesene formation during oolong tea processing [J]. Food Industry Science and Technology, 2020,41(15): 135-142.

[62]    Chang Y N, Zhu C, Jiang J, et al. Epigenetic regulation in plant abiotic stress responses [J]. Journal of Integrative Plant Biology, 2020, 62(5): 563-580.

[63]    Saini P, Bhatia S, Mahajan M, et al. ELONGATED HYPOCOTYL5 negatively regulates DECREASE WAX BIOSYNTHESIS to increase survival during UV-B stress [J]. Plant Physiology, 2020, 184(4): 2091-2106.

[64]    Chen S, Wang P J, Kong W L, et al. Gene mining and genomics-assisted breeding empowered by the pangenome of tea plant Camellia sinensis [J]. Nature Plants, 2023, 9(12): 1986-1999.

[65]    Pak S, Li C H. Progress and challenges in applying CRISPR/Cas techniques to the genome editing of trees [J]. Forestry Research, 2022, 22(1): 46-59.

[66]    Alagarsamy K, Shamala L F, Wei S. Protocol: high-efficiency in-planta Agrobacterium-mediated transgenic hairy root induction of Camellia sinensis var. sinensis [J]. Plant Methods, 2018, 14(1): 17. https://doi.org/10.1186/s13007-018-0285-8.

[67]    Mao Y L, Li H, Wang Y, et al. Rapid monitoring of tea plants under cold stress based on UAV multi-sensor data [J]. Computers and Electronics in Agriculture, 2023, 213: 108176. https://doi.org/10.1016/j.compag.2023.108176.

[68]    Sun W, Yuan H Z, Lei T Y, et al. First report of anthracnose caused by Colletotrichum kahawae and Colletotrichum horri on tea-oil tree in China [J]. Plant Disease, 2022, 107(6): 1944. https://doi.org/10.1094/PDIS-06-22-1312-PDN.

[69]    Zhang L, Wang F P, Li Q, et al. Population structure and genetic differentiation of tea green leafhopper, Empoasca (Matsumurasca) onukii, in China based on microsatellite markers [J]. Scientific Reports, 2019, 9: 1202. https://doi.org/10.1038/s41598-018-37881-0.

[70]    Tao J, Xie Y J, Wang W F, et al. Elevational gradient of climate-driving effects on cropland ecosystem net primary productivity in alpine region of Southwest China [J]. Remote Sensing, 2022, 14(13): 3069-3081.

[71]    Gu D C, Wu S H, Wang Y X, et al. Tea green leafhopper infestations affect tea plant growth by altering the synthesis of brassinolide [J]. Plant, Cell & Environment, 2024, 47(10): 3780-3796.

[72]    Duan Y, Wang G, Liang L Y, et al. Intercropping fruit trees in tea plantation improves soil properties and the formation of tea quality components [J]. Plant Physiology and Biochemistry, 2024, 210(4): 108574-108586.

[73]    Sun L T, Xue D, Wang Y, et al. Tea-soybean intercropping improves tea quality and nutrition uptake by inducing changes of rhizosphere bacterial communities [J]. Microorganisms, 2022, 10(11): 2149-2160.

[74]    Zhang G L, Chu X B, Zhu H Y, et al. The response of soil nutrients and microbial community structures in long-term tea plantations and diverse agroforestry intercropping systems [J]. Sustainability, 2021, 13(14): 7799. https://doi.org/10.3390/su13147799.

[75]    Lin J, Lin H, Li C, et al. Unveiling characteristic metabolic accumulation over enzymatic-catalyzed process of Tieguanyin oolong tea manufacturing by DESI-MSI and multiple-omics [J]. Food Research International, 2024, 181(4): 114136-114151.

[76]    Wu L Y, Chen X L, Lin J Q, et al. Study on dynamic alterations of volatile organic compounds reveals aroma development over enzymatic-catalyzed process of Tieguanyin oolong tea production [J]. Food Chemistry: Molecular Sciences, 2024, 9(16): 100227. https://doi.org/10.1016/j.fochms.2024.100227.

[77]    Zhang G Y, Wei J J, Li L Y, et al. Lipidomics, transcription analysis, and hormone profiling unveil the role of CsLOX6 in MeJA biosynthesis during black tea processing [J]. Horticulture Research, 2024, 11(3): uhae032. https://doi.org/10.1093/hr/uhae032.

[78]    Luo Z W, Zhang Q, Yang Y W, et al. Effect of rainy-day harvesting on the aroma profile of Wuyi rock tea based on GC-MS and chemical analysis [J]. Food Chemistry: X, 2025, 29(8): 102753. https://doi.org/10.1016/j.fochx.2025.102753.

[79]    Li Q M, He J H, Yu X R, et al. Exploring the taste code of light-withered tea: the mechanism of nonvolatile metabolite changes in LED light-withered oolong tea (Camellia sinensis) [J]. Food Chemistry: X, 2025, 30(10): 102873. https://doi.org/10.1016/j.fochx.2025.102873.

[80]    Chen Q, Chen H Y, Qu H, et al. Improper use of spinetoram on tea plants can inhibit the predation ability of Arma chinensis [J]. Ecotoxicology and Environmental Safety, 2025, 302(1): 118638. https://doi.org/10.1016/j.ecoenv.2025.118638.

[81]    Wang L Y, Sun P J, Huang M X, et al. Impact of kombucha fermentation on the flavor and physicochemical properties of tea (Camellia sinensis) flower infusions [J]. Food Frontiers, 2025, 6(2): 750-762

[82]    Orita T, Chogahara S, Okuda M, et al. Extraction efficiency and alpha-glucosidase inhibitory activities of green tea catechins by different infusion methods [J]. Foods, 2023, 12(13): 2611. https://doi.org/10.3390/foods12132611.

[83]    Serdar G, Demir E, Skmen M. Sequential green extraction of caffeine and catechins from green tea [J]. International Journal of Secondary Metabolite, 2019, 6(3): 283-291.

[84]    Zhang X L, Lu M R, Li M, et al. Efficient callus induction and regeneration of tea plant (camellia sinensis) [J]. Scientific Reports, 2025, 15: 26848. https://doi.org/10.1038/s41598-025-12271-5.

[85]    Wang Y, Kan Z, Thompson H J, et al. Impact of six typical processing methods on the chemical composition of tea leaves using a single Camellia sinensis cultivar, Longjing 43 [J]. Journal of Agricultural and Food Chemistry, 2019, 67(19): 5423-5436.

[86]    Xie G H, Yan J N, Lu A X, et al. Characterizing relationship between chemicals and in vitro bioactivities of teas made by six typical processing methods using a single Camellia sinensis cultivar, Meizhan [J]. Bioengineered, 2021, 12(1): 1251-1263.

[87]    Niu Z R, Zhang Y, Han J C, et al. The soil ecological stoichiometry characteristics of the highest latitude areas in the main tea-producing regions of China [J]. Agronomy, 2024, 14(7): 1359. https://doi.org/10.3390/agronomy14071359.

[88]    Wang M, Yang J, Li J L, et al. Effects of temperature and light on quality-related metabolites in tea [Camellia sinensis (L.) Kuntze] leaves [J]. Food Research International, 2022, 161: 111882. https://doi.org/10.1016/j.foodres.2022.111882.

[89]    Liu D D, Zhang C Y, Ye Y Y, et al. TEA5K: a high-resolution and liquid-phase multiple-SNP array for molecular breeding in tea plant [J]. Journal of Nanobiotechnology, 2025, 23(1): 481. https://doi.org/10.1186/s12951-025-03533-5.

[90]    Tao L L, Ting Y J, Chen H R, et al. Core collection construction of tea plant germplasm in Anhui Province based on genetic diversity analysis using simple sequence repeat markers [J]. Journal of Integrative Agriculture, 2023, 22(9): 2719-2728.

[91]    Yadav S, Rahim M S, Devi A, et al. Revolutionizing speciality teas: multi-omics prospective to breed anthocyanin-rich tea [J]. Food Research International, 2025, 209(5): 116312. https://doi.org/10.1016/j.foodres.2025.116312.

[92]    Anastasiya E, Ivan F, Anastasia F, et al. Targeted genome modification in protoplasts of a tea cultivar Kolkhida using RNA-guided Cas9 endonuclease [J]. AoB Plants, 2025, 17(5): plaf056. https://doi.org/10.1093/aobpla/plaf056.