[1] Li T T, Wang S R, Shi D D, et al. Phosphate deficiency induced by infection promotes synthesis of anthracnose-resistant anthocyanin-3-O-galactoside phytoalexins in the Camellia sinensis plant[J]. Horticulture Research, 2023, 10(12): uhad222. https://doi.org/10.1093/hr/uhad222.
[2] Wang L, Wang Y C, Cao H L, et al. Transcriptome analysis of an anthracnose-resistant tea plant cultivar reveals genes associated with resistance to Colletotrichum camelliae[J]. PLoS One, 2016, 11(2): e0148535. https://doi.org/10.1371/journal.pone.0148535.
[3] 韩雨欣. 白化茶树品种的代谢差异及其对茶炭疽菌的抗性研究[D]. 重庆: 西南大学, 2024: 10-64.
Han Y X.Metabolic differences of albino tea cultivars and their resistance to Colletotrichum camelliae[D]. Chongqing: Southwest University, 2024: 10-64.
[4] Lü W Y, Xu Y J, Jiang H, et al. An NBS-LRR-encoding gene CsRPM1 confers resistance to the fungus Colletotrichum camelliae in tea plant[J]. Beverage Plant Research, 2023, 3: 13. https://doi.org/10.48130/BPR-2023-0013
[5] Zhang C Y, Li H L, Mei P, et al.QTL detection and candidate gene analysis of the anthracnose resistance locus in tea plant (Camellia sinensis)[J]. Journal of Integrative Agriculture, 2025, 24(6): 2240-2250.
[6] 贡长怡, 刘姣姣, 邓强, 等. 茶树炭疽病病原菌鉴定及其致病性分析[J]. 园艺学报, 2022, 49(5): 1092-1101.
Gong C Y, Liu J J, Deng Q, et al.Identification and pathogenicity analysis of anthracnose pathogen in tea plant[J]. Acta Horticulturae Sinica, 2022, 49(5): 1092-1101.
[7] Jeyaraj A, Elango T, Chen X, et al.Advances in understanding the mechanism of resistance to anthracnose and induced defence response in tea plants[J]. Molecular Plant Pathology, 2023, 24(10): 1330-1346.
[8] 石媛, 王昆, 田家顺. 茶树炭疽病防治田间药效试验[J]. 湖南农业科学, 2023(4): 73-75. https://doi.org/10.16498/j.cnki.hnnykx.2023.004.015.
Shi Y, Wang K, Tian J S.Field efficacy test for control of tea anthracnose[J]. Hunan Agricultural Sciences, 2023(4): 73-75. https://doi.org/10.16498/j.cnki.hnnykx.2023.004.015.
[9] 唐朝阳, 孔丽娅, 胡骞, 等. 贝莱斯芽孢杆菌YJK1鉴定及其对茶炭疽病的拮抗效果[J]. 茶叶科学, 2024, 44(3): 443-452.
Tang C Y, Kong L Y, Hu Q, et al.Identification of Bacillus velezensis YJK1 and its antagonistic effect on tea anthracnose[J]. Journal of Tea Science, 2024, 44(3): 443-452.
[10] Peng X J, Wang Q C, Zhang S K, et al.Colletotrichum species associated with Camellia anthracnose in China[J]. Mycosphere, 2023, 14(2): 130-157.
[11] Tao L L, Zhu J Y, Hu J B, et al. Pangenome analyses of tea plants reveal structural variations driving gene expression alterations and agronomic trait diversification[J]. Nature Communications, 2026, 17: 372. https://doi.org/10.1038/s41467-025-67060-5.
[12] Xia E H, Tong W, Wu Q, et al. Tea plant genomics: achievements, challenges and perspectives[J]. Horticulture Research, 2020, 7(1): 7. https://doi.org/10.1038/s41438-019-0225-4.
[13] Dean R, Van Kan J A L, Pretorius Z A, et al. The top 10 fungal pathogens in molecular plant pathology[J]. Molecular Plant Pathology, 2012, 13(4): 414-430.
[14] Cannon P F, Damm U, Johnston P R, et al.Colletotrichum-current status and future directions[J]. Studies in Mycology, 2012, 73(1): 181-213.
[15] Von Arx J A. Die Arten der Gattung Colletotrichum Cda[J]. Phytopathologische Zeitschrift, 1957, 29(4): 413-468.
[16] Baxter A P, Van der Westhuizen G C A, Eicker A. Morphology and taxonomy of South African isolates of Colletotrichum[J]. South African Journal of Botany, 1983, 2(4): 259-289.
[17] 王玉春. 中国茶树炭疽菌系统发育学研究及茶树咖啡碱抗炭疽病的作用[D]. 杨凌: 西北农林科技大学, 2016: 17-43.
Wang Y C.Phylogenetic study of Colletotrichum from tea plants in China and the role of caffeine in resistance to anthracnose[D]. Yangling: Northwest A & F University, 2016: 17-43.
[18] Lin S R, Yu S Y, Chang T D, et al. First report of anthracnose caused by Colletotrichum fructicola on tea in Taiwan[J]. Plant Disease, 2021, 105: 710. https://doi.org/10.1094/PDIS-06-20-1288-PDN.
[19] Yu L, Lan G B, Yang Y G, et al.First report of anthracnose caused by Colletotrichum fructicola on Brassica parachinensis in China[J]. Crop Protection, 2022, 154: 105842. https://doi.org/10.1016/J.CROPRO.2021.105842.
[20] Fernández R L, Rivera M C, Varsallona B, et al.Disease prevalence and symptoms caused by Alternaria tenuissima and Pestalotiopsis guepinii on blueberry in Entre Ríos and Buenos Aires, Argentina[J]. American Journal of Plant Sciences, 2015, 6(19): 3082-3090.
[21] 刘威. 茶树炭疽病的病原鉴定及其遗传多样性分析[D]. 福州: 福建农林大学, 2013: 9-33.
Liu W.Pathogen identification and genetic diversity analysis of tea anthracnose[D]. Fuzhou: Fujian Agriculture and Forestry University, 2013: 9-33.
[22] Lu Q H, Wang Y C, Li N N, et al.Differences in the characteristics and pathogenicity of Colletotrichum camelliae and C. fructicola isolated from the tea plant[Camellia sinensis (L.) O. Kuntze] [J]. Frontiers in Microbiology, 2018, 9: 3060. https://doi.org/10.3389/fmicb.2018.03060.
[23] Liu F, Weir B S, Damm U, et al. Unravelling Colletotrichum species associated with Camellia: employing ApMat and GS loci to resolve species in the C. gloeosporioides complex[J]. Persoonia, 2015, 35: 63-86. https://doi.org/10.3767/003158515X687597.
[24] 邵鑫. 茶炭疽病的发生及其防治[J]. 蚕桑茶叶通讯, 2001(1): 14.
Shao X.Occurrence and control of tea anthracnose[J]. Newsletter of Sericulture and Tea, 2001(1): 14.
[25] 邹雨伽, 杨柳, 张秀琼, 等. 基于GIS的茶叶生产适宜性区划研究——以旺苍县为例[J]. 西南大学学报(自然科学版), 2021, 43(3): 53-59.
Zou Y J, Yang L, Zhang X Q, et al.GIS-based suitability regionalization for tea production: a case study of Wangcang County[J]. Journal of Southwest University (Natural Science Edition), 2021, 43(3): 53-59.
[26] 林德锋, 吴顺章, 黄大龙, 等. 漳州铁观音茶炭疽病发生规律及化学防治试验[J]. 浙江农业科学, 2012(8): 1169-1171.
Lin D F, Wu S Z, Huang D L, et al.Occurrence regularity and chemical control experiment of Tieguanyin tea anthracnose in Zhangzhou[J]. Journal of Zhejiang Agricultural Sciences, 2012(8): 1169-1171.
[27] 唐美君, 郭华伟, 姚惠明, 等. 龙井茶区茶炭疽病的发生规律[J]. 浙江农业科学, 2019, 60(10): 1763-1765.
Tang M J, Guo H W, Yao H M, et al.Occurrence regularity of tea anthracnose in Longjing tea area[J]. Journal of Zhejiang Agricultural Sciences, 2019, 60(10): 1763-1765.
[28] 张新春, 彭元科, 王家保. 不同来源荔枝胶孢炭疽菌生物学特性研究[J]. 中国果树, 2015(3): 27-31.
Zhang X C, Peng Y K, Wang J B.Biological characteristics of Colletotrichum gloeosporioides from different sources of Litchi[J]. China Fruits, 2015(3): 27-31.
[29] 刘芸芸, 丁一汇. 2020年超强梅雨特征及其成因分析[J]. 气象, 2020, 46(11): 1393-1404.
Liu Y Y, Ding Y H.Characteristics and causes of the 2020 super Meiyu[J]. Meteorological Monthly, 2020, 46(11): 1393-1404.
[30] 潘蔚娟, 蒋承霖. 华南地区春雨进程的气候特点及其气候季节内振荡[J]. 热带气象学报, 2014, 30(1): 83-91.
Pan W J, Jiang C L.Climatic characteristics and intra-seasonal oscillation of spring rainfall process in South China[J]. Journal of Tropical Meteorology, 2014, 30(1): 83-91.
[31] 汪孝科. 茶叶炭疽病的防治试验[J]. 农业与技术, 2014, 34(6): 130-131.
Wang X K.Control experiment of tea anthracnose[J]. Agriculture and Technology, 2014, 34(6): 130-131.
[32] 王兴娥, 马媛, 孙友慧, 等. 都匀市螺丝壳秋季茶园主要真菌病害调查及防治建议[J]. 农业与技术, 2024, 44(8): 14-17.
Wang X E, Ma Y, Sun Y H, et al.Investigation and control suggestions of main fungal diseases in autumn tea garden of Luosike, Duyun City[J]. Agriculture and Technology, 2024, 44(8): 14-17.
[33] 李志伟, 谭玉梅, 任锡毅, 等. 茶树炭疽病病原菌的绿色荧光蛋白基因标记及其侵染研究[J]. 基因组学与应用生物学, 2020, 39(8): 3510-3518.
Li Z W, Tan Y M, Ren X Y, et al.GFP gene labeling and infection study of tea anthracnose pathogen[J]. Genomics and Applied Biology, 2020, 39(8): 3510-3518.
[34] Li M, Liu J, Zhou G.Histopathological and ultrastructural observations of Camellia oleifera infected with Colletotrichum fructicola[J]. Australasian Plant Pathology, 2021, 50(5): 523-531.
[35] Pandey A K, Sinniah G D, Babu A, et al.How the global tea industry copes with fungal diseases: challenges and opportunities[J]. Plant Disease, 2021, 105(7): 1868-1879.
[36] Lu Q H, Wang Y C, Xiong F, et al. Integrated transcriptomic and metabolomic analyses reveal the effects of callose deposition and multihormone signal transduction pathways on the tea plant-Colletotrichum camelliae interaction[J]. Scientific Reports, 2020, 10(1): 12858. https://doi.org/10.1038/s41598-020-69729-x.
[37] Li Y X, Lin K Q, Wang A R, et al.Multi-omics analysis of molecular mechanisms driving the grafting-enhanced resistance of tea plants to Colletotrichum camelliae[J]. Frontiers in Plant Science, 2025, 16: 1750493. https://doi.org/10.3389/FPLS.2025.1750493.
[38] Liu W, Guo X Y, Chen Q, et al.Brassinolide-induced resistance enhances antioxidant defense and metabolic pathways against anthracnose in Camellia sinensis ‘Fuding Dabaicha’[J]. Scientia Horticulturae, 2025, 349: 114258. https://doi.org/10.1016/j.scienta.2025.114258.
[39] Qiao X Y, Zhang S H, He S N, et al.Urate oxidase treatment increases the quality of autumn yellowish ‘Yinghong 9’ black tea[J]. LWT, 2023, 184: 115092. https://doi.org/10.1016/j.lwt.2023.115092.
[40] 黄镇雄, 吴家豪, 叶思潭, 等. 基于高光谱成像技术的茶树炭疽病早期诊断方法[J]. 农业工程学报, 2025, 41(23): 193-205.
Huang Z X, Wu J H, Ye S T, et al.Early diagnosis method of tea anthracnose based on hyperspectral imaging technology[J]. Transactions of the Chinese Society of Agricultural Engineering, 2025, 41(23): 193-205.
[41] Zrimec J, Correa S, Zagorščak M, et al. Evaluating plant growth-defense trade-offs by modeling the interaction between primary and secondary metabolism[J]. Proceedings of the National Academy of Sciences, 2025, 122(32): e2502160122. https://doi.org/10.1073/pnas.2502160122.
[42] Xu L Y, Su J J, Zhang C K, et al. Identification of key genes associated with anthracnose resistance in Camellia sinensis[J]. PLoS One, 2025, 20(6): e0326325. https://doi.org/10.1371/JOURNAL.PONE.0326325.
[43] 张景贺, 张瑾, 孙晓玲, 等. 10种化学杀菌剂对茶树炭疽病菌(Colletotrichum camelliae)的室内毒力测定[J]. 中国茶叶, 2024, 46(8): 35-39.
Zhang J H, Zhang J, Sun X L, et al.Indoor toxicity determination of 10 chemical fungicides against tea anthracnose pathogen (Colletotrichum camelliae)[J]. China Tea, 2024, 46(8): 35-39.
[44] 石文波, 黄楚平, 陈宇航, 等. 纳米氢氧化铜对茶炭疽病菌的抑制活性及作用机制[J]. 农药学学报, 2024, 26(5): 962-973.
Shi W B, Huang C P, Chen Y H, et al.Inhibitory activity and mechanism of nano-copper hydroxide against tea anthracnose pathogen[J]. Chinese Journal of Pesticide Science, 2024, 26(5): 962-973.
[45] 吴庆丽, 秦刚, 李慧, 等. 助剂激健与杀菌剂混用对3种茶树病害的防效[J]. 中国植保导刊, 2020, 40(8): 69-71, 77.
Wu Q L, Qin G, Li H, et al.Control efficacy of adjuvant Jijian mixed with fungicides on three tea diseases[J]. China Plant Protection, 2020, 40(8): 69-71, 77.
[46] Liu L, Guan H N, Jiao M M, et al. Identification of two antagonistic fungi and antifungal activity analysis against anthracnose in tea plant (Camellia sinensis)[J]. Beverage Plant Research, 2024, 4: e032. https://doi.org/10.48130/bpr-0024-0020.
[47] 刘辉, 冯月玲, 周罗娜, 等. 茶炭疽病拮抗木霉菌株的分离鉴定及其对茶炭疽病的拮抗作用[J]. 西南农业学报, 2025, 38(4): 757-768.
Liu H, Feng Y L, Zhou L N, et al.Isolation and identification of antagonistic Trichoderma strains against tea anthracnose and their antagonistic effects[J]. Southwest China Journal of Agricultural Sciences, 2025, 38(4): 757-768.
[48] 尤佳琪, 杜然, 顾卫红, 等. 拟康宁木霉T-51菌株生物学特性及其生物防治潜力[J]. 植物保护学报, 2022, 49(3): 946-955.
You J Q, Du R, Gu W H, et al.Biological characteristics and biocontrol potential of Trichoderma koningiopsis T-51 strain[J]. Journal of Plant Protection, 2022, 49(3): 946-955.
[49] Cortez-Lázaro A A, Chavez-Castillo J I, Romero-Bozzetta J L, et al. From mechanisms to networks: a global bibliometric and functional synthesis of Trichoderma in plant disease management[J]. Journal of Agriculture and Food Research, 2026, 26: 102706. https://doi.org/10.1016/j.jafr.2026.102706.
[50] Chen Q X, Song Y J, An Y X, et al. Mechanisms and impact of rhizosphere microbial metabolites on crop health, traits, functional components: a comprehensive review[J]. Molecules, 2024, 29(24): 5922. https://doi.org/10.3390/molecules29245922.
[51] Bardas G A, Lagopodi A L, Kadoglidou K, et al.Biological control of three Colletotrichum lindemuthianum races using Pseudomonas chlororaphis PCL1391 and Pseudomonas fluorescens WCS365[J]. Biological Control, 2009, 49(2): 139-145.
[52] 宫安东, 韩萌真, 孔宪巍, 等. 茶树内生菌的应用性研究进展[J]. 信阳师范学院学报(自然科学版), 2017, 30(1): 168-172.
Gong A D, Han M Z, Kong X W, et al.Research progress on application of tea endophytes[J]. Journal of Xinyang Normal University (Natural Science Edition), 2017, 30(1): 168-172.
[53] 施云龙. 茶树抗炭疽病和抗冻机制及评价研究[D]. 杭州: 浙江大学, 2020: 50-84.
Shi Y L.Study on resistance mechanism to anthracnose and freezing and evaluation in tea plant[D]. Hangzhou: Zhejiang University, 2020: 50-84.
[54] 孙春霞, 邵元海, 周红, 等. 茶树六种重要叶部病害研究进展[J]. 茶叶, 2020, 46(2): 71-76.
Sun C X, Shao Y H, Zhou H, et al.Research progress on six important leaf diseases of tea plant[J]. Journal of Tea, 2020, 46(2): 71-76.
[55] 刘荣, 周爽爽, 韦玲冬, 等. 茶炭疽病菌的生物学特性研究[J]. 福建茶叶, 2020, 42(11): 13-14.
Liu R, Zhou S S, Wei L D, et al.Study on biological characteristics of tea anthracnose pathogen[J]. Tea in Fujian, 2020, 42(11): 13-14.
[56] 唐美君, 郭华伟, 石碧鹏, 等. 新昌雪溪茶场茶炭疽病发生动态与绿色防控示范[J]. 中国茶叶, 2022, 44(12): 29-33.
Tang M J, Guo H W, Shi B P, et al.Occurrence dynamics and green control demonstration of tea anthracnose in Xinchang Xuexi Tea Farm[J]. China Tea, 2022, 44(12): 29-33.
[57] 向雪琴. 茶叶病虫害种类及其生态防控策略[J]. 贵茶, 2024(6): 21-24.
Xiang X Q.Types of tea diseases and pests and their ecological control strategies[J]. Guizhou Tea, 2024(6): 21-24.
[58] 周凌云, 李静, 王沅江. 茶炭疽病菌的分离、培养及PCR检测[J]. 福建茶叶, 2009, 31(3): 11-12.
Zhou L Y, Li J, Wang Y J.Isolation, culture and PCR detection of tea anthracnose pathogen[J]. Tea in Fujian, 2009, 31(3): 11-12.
[59] 谢宝林. 茶炭疽病的识别与防治[J]. 农技服务, 2007(2): 63.
Xie B L.Identification and control of tea anthracnose[J]. Agricultural Technology Service, 2007(2): 63.
[60] Takeda Y, et al. Breeding of the early budding cultivar ‘Saemidori’ and the late budding cultivar ‘Okumidori’[J]. Breeding Science, 2006, 8: 113-117. https://doi.org/10.1270/jsbbr.8.113.
[61] Han R, Mei H L, Huang Q W, et al. CsNAC17 enhances resistance to Colletotrichum gloeosporioides by interacting with CsbHLH62 in Camellia sinensis[J]. Horticulture Research, 2025, 12(2): uhae295. https://doi.org/10.1093/hr/uhae295.
[62] Li C Y, Wu F X, Yang L, et al. UGT74B5-mediated glucosylation at ortho hydroxyl groups of benzoic acid derivatives regulating plant immunity to anthracnose in tea plants[J]. Horticulture Research, 2025, 12(4): uhaf009. https://doi.org/10.1093/hr/uhaf009.
[63] Tao Y N, Wang P K, Gong Y L, et al.A positive regulator CsPR10-9 confers resistance to anthracnose (Colletotrichum gloeosporioides) is negatively regulated by CsMYB72 in tea plants[J]. Plant, Cell & Environment, 2025, 48(9): 6965-6981.
[64] Jiang T, Li X, Song J M, et al. Long non-coding RNA Cslnc256 regulates tea plant resistance to anthracnose by suppressing CsmiR395-mediated sulfate metabolism[J]. The Plant Journal, 2026, 125(4): e70720. https://doi.org/10.1111/tpj.70720.
[65] 梁俊杰, 王东霞. 基于改进EfficientNet-Lite适用于复杂背景下茶树病害识别轻量级移动模型[J]. 农业技术与装备, 2024(9): 23-25, 28.
Liang J J, Wang D X.Lightweight mobile model for tea disease recognition in complex background based on improved EfficientNet-Lite[J]. Agricultural Technology and Equipment, 2024(9): 23-25, 28.
[66] 宋军, 张佑丞, 徐锋, 等. 基于改进DETR模型的轻量化茶叶病虫害检测方法[J]. 实验室研究与探索, 2025, 44(8): 39-47, 54.
Song J, Zhang Y C, Xu F, et al.Lightweight tea pest and disease detection method based on improved DETR model[J]. Research and Exploration in Laboratory, 2025, 44(8): 39-47, 54.
[67] 王兴娥, 刘荣, 杨欣, 等. 茶炭疽病的研究进展[J]. 湖北植保, 2024(2): 13-17.
Wang X E, Liu R, Yang X, et al.Research progress on tea anthracnose[J]. Hubei Plant Protection, 2024(2): 13-17.
[68] 王玉春, 刘守安, 卢秦华, 等. 中国茶树炭疽菌属病害研究进展及展望[J]. 植物保护学报, 2019, 46(5): 954-963.
Wang Y C, Liu S A, Lu Q H, et al.Research progress and prospects of Colletotrichum diseases of tea plant in China[J]. Journal of Plant Protection, 2019, 46(5): 954-963.
[69] Fan K, Zhang J, Wang M, et al.Development and application of SNP-KASP markers based on genes related to nitrogen uptake, assimilation and allocation in tea plant (Camellia sinensis L.)[J]. Agronomy, 2022, 12(10): 2534-2534.