LIU Guanhua1 , 2 , 3 , YANG Mei1 , 2 , 3 , FU Jianyu1 , 2* . 茶树胡萝卜素合成关键基因CsLCYb 和CsLCYe 的克隆与功能鉴定[J]. 茶叶科学, 2019 , 39(3) : 257 -266 . DOI: 10.13305/j.cnki.jts.2019.03.003

[1] Tanaka Y, Sasaki N, Ohmiya A. Biosynthesis of plant pigments: anthocyanins, betalains and carotenoids [J]. The Plant Journal, 2008, 54(4): 733-749. [2] Nishio J N. Why are higher plants green？Evolution of the higher plant photosynthetic pigment complement [J]. Plant Cell & Environment, 2010, 23(6): 539-548. [3] Grotewold E. The genetics and biochemistry of floral pigments [J]. Annual Review of Plant Biology, 2006, 57(1): 761-780. [4] Rockwell N C, Su Y S, Lagarias J C. Phytochrome structure and signaling mechanisms [J]. Annual Review of Plant Biology, 2006, 57(1): 837-858. [5] Frank H A, Cogdell R J. Carotenoids in photosynthesis [J]. Photochem Photobiol, 1996, 63(3): 257-264. [6] Lintig J V. Colors with functions: elucidating the biochemical and molecular basis of carotenoid metabolism [J]. Annual Review of Nutrition, 2010, 30(1): 35-56. [7] Nambara E, Marion-Poll A. Abscisic acid biosynthesis and catabolism [J]. Annual Review of Plant Biology, 2005, 56: 165-185. [8] Al-Babil S, Bouwmeester H J. Strigolactones, a novel carotenoid-derived plant hormone [J]. Annual Review of Plant Biology, 2015, 66(1): 161-186. [9] Tanaka R, Tanaka A. Tetrapyrrole biosynthesis in higher plants [J]. Annual Review of Plant Biology, 2007, 58: 321-346. [10] Bach T J. Some new aspects of isoprenoid biosynthesis in plants—a review [J]. Lipids, 1995, 30(3): 191-202. [11] McGarvey D J, Croteau R. Terpenoid metabolism [J]. Plant Cell, 1995, 7(7): 1015-1026. [12] 李莉, 高凌云, 董越, 等. 植物类异戊二烯生物合成相关酶基因研究进展[J]. 浙江师范大学学报(自然科学版), 2008, 31(4): 461-466. [13] 高慧君, 明家琪, 张雅娟, 等. 园艺植物中类胡萝卜素合成与调控的研究进展[J]. 园艺学报, 2015, 42(9): 1633-1648. [14] Cheng A, Lou Y, Mao Y, et al. Plant terpenoids: biosynthesis and ecological functions [J]. Journal of Integrative Plant Biology, 2007, 49(2): 179-186. [15] Cunningham F X Jr, Gantt E. Genes and enzymes of carotenoid biosynthesis in plants [J]. Annual Review of Plant Physiology and Plant Molecular Biology, 1998, 49(1): 557-583. [16] Takaichi S. Carotenoids in algae: distributions, biosyntheses and functions [J]. Marine Drugs, 2011, 9(6): 1101-1118. [17] Takaichi S. Tetraterpenes: carotenoids [M]∥Merillon J M, Ramawat K G. Berlin: Springer: Natural Products, 2013: 3251-3283. [18] Schnurr G, Misawa N, Sandmann G. Expression, purification and properties of lycopene cyclase from Erwinia uredovora [J]. Biochemical Journal, 1996, 315(3): 869-874. [19] Hisashi H, Norihiko M. Novel approaches and achievements in biosynthesis of functional isoprenoids in Escherichia coli [J]. Applied Microbiology & Biotechnology, 2009, 84(6): 1021-1031. [20] Sugiyama K, Ebisawa M, Yamada M, et al. Functional lycopene cyclase (crua) in cyanobacterium, arthrospira platensis nies-39, and its role in carotenoid synthesis [J]. Plant & Cell Physiology, 2017, 58(4): 831-838. [21] 宛晓春. 茶叶生物化学[M]. 3版. 北京: 中国农业出版社, 2003, 23-29. [22] Wang L, Yue C, Cao H, et al. Biochemical and transcriptome analyses of a novel chlorophyll-deficient chlorina tea plant cultivar [J]. BMC Plant Biology, 2014, 14: 352. doi.org/10.1186/s12870-014-0352-x. [23] Li N, Yang Y, Ye J, et al. Effects of sunlight on gene expression and chemical composition of light-sensitive albino tea plant [J]. Plant Growth Regulation, 2016, 78(2): 253-262. [24] 邓静, 王远兴, 丁建. 白茶与安吉白茶的研究进展[J]. 食品工业科技, 2013, 34(4): 369-374. [25] Grabherr M G, Haas B J, Yassour M, et al. Full-length transcriptome assembly from RNA-Seq data without a reference genome [J]. Nature Biotechnology, 2011, 29(7): 644-652. [26] Cunningham F X Jr, Sun Z, Chamovitz D, et al. Molecular structure and enzymatic function of lycopene cyclase from the cyanobacterium synechococcus sp. strain PCC7942 [J]. Plant Cell, 1994, 6(8): 1107-1121. [27] Francis X C, Barry P, Zairen S, et al. Functional analysis of the beta and epsilon lycopene cyclase enzymes of Arabidopsis reveals a mechanism for control of cyclic carotenoid formation [J]. Plant Cell, 1996, 8(9): 1613-1626. [28] 金应福, 韩莉, 张莎莎, 等. 通过番茄红素环化酶的优化构建β-胡萝卜素高产菌株[J]. 生物工程学报, 2017, 33(11): 1814-1826. [29] Gasteiger E, Hoogland C, Gattiker A, et al. Protein identification and analysis tools on the ExPASy Server [M]∥John M Walker. The proteomics protocols handbook. humana press, 2005: 571-607. [30] Kumar S, Stecher G, Tamura K. MEGA7: molecular evolutionary genetics analysis version 7.0 for bigger datasets [J]. Molecular Biology and Evolution, 2016, 33(7): 1870-1874. [31] Wei C, Yang H, Wang S, et al. Draft genome sequence of Camellia sinensis var. sinensis provides insights into the evolution of the tea genome and tea quality [J]. Proc Natl Acad Sci U S A, 2018, 115(18): E4151-E4158. [32] 虞富莲. 中黄1号、中黄2号的特异性、一致性和稳定性[J]. 中国茶叶, 2016, 38(3): 14-16. [33] Pecker I, Gabbay R, Cunningham F X Jr, et al. Cloning and characterization of the cDNA for lycopene β-cyclase from tomato reveals decrease in its expression during fruit ripening [J]. Plant Molecular Biology, 1996, 30(4): 807-819. [34] Pogson B, McDonald K A, Truong M, et al. Arabidopsis carotenoid mutants demonstrate that lutein is not essential for photosynthesis in higher plants [J]. Plant Cell, 1996, 8(9): 1627-1639. [35] 朱跃辉, 姜建国. 类胡萝卜素代谢途径中相关番茄红素环化酶的功能[J]. 中国食品添加剂, 2005(5): 40-43. [36] Ronen G, Cohen M, Zamir D, et al. Regulation of carotenoid biosynthesis during tomato fruit development: expression of the gene for lycopene epsilon cyclase is down-regulated during ripening and is elevated in the mutant delta [J]. Plant Journal, 1999, 17(4): 341-351. [37] Norman I K, Elizabeth J J. Carotenoid actions and their relation to health and disease [J]. Molecular Aspects of Medicine, 2005, 26(6): 459-516. [38] Feng L, Gao M J, Hou R Y, et al. Determination of quality constituents in the young leaves of albino tea cultivars [J]. Food Chemistry, 2014, 155(11): 98-104.